Guadayol, O., Schuech, R. & Humphries, S. (2023) Optimal cell length for exploration and exploitation in chemotactic planktonic bacteria. bioRxiv doi:10.1101/2023.10.05.560636.

Soulsbury, C.D. & Humphries, S. (2022). Biophysical determinants and constraints on sperm swimming velocity. Cells 11, 3360.

Guadayol, Ò., Mendonca, T., Segura-Noguera, M., Wright, A. J., Tassieri, M. & Humphries, S. (2020). Microrheology reveals microscale viscosity gradients in planktonic systems. PNAS , 118, e2011389118.

Venditti, C., Baker, J., Meade, A., Benton, M. & Humphries, S. (2020). 150 million years of sustained increase in pterosaur flight efficiency. Nature, 587, 83–86

Walker, M., Hammel, J. U., Wilde, F., Hoehfurtner, T., Humphries, S. & Schuech, R. (2020). Estimation of sinking velocities using free-falling dynamically scaled models: foraminifera as a test case. J Exp Biol, jeb.230961 doi: 10.1242/jeb.230961

El baidouri, F., Venditti, C., Suzuki, S., Meade, A. & Humphries, S. (2020). Phenotypic reconstruction of the last universal common ancestor reveals a complex cell. bioRxiv

Baker, J., Humphries, S., Ferguson-Gow, H., Meade, A. & Venditti, C. (2020) Rapid decreases in relative testes mass among monogamous birds but not in other vertebrates. Ecology Letters, 23, 283-192, doi: 10.1111/ele.13431

Schuech, R., Hoehfurtner, T., Smith, D. & Humphries, S. (2019) Motile curved bacteria are Pareto-optimal. PNAS, 726, 201818997,

Scott, G.W., Humphries, S. & Henri, D.C. (2019). Expectation, motivation, engagement and ownership: using student reflections in the conative and affective domains to enhance residential field courses. Journal of Geography in Higher Education, 43(3), pp.280–298.

Walker, M.W. & Humphries, S. (2019) 3D Printing: Applications in evolution and ecology. Ecol & Evoln

Guadayol, O., Thornton, K.L. & Humphries, S. (2017) Cell morphology governs directional control in swimming bacteria. Sci Rep 7(1):2061

El Baidouri, F., Venditti, C. & Humphries, S. (2016) Independent evolution of shape and motility allows evolutionary flexibility in Firmicutes bacteria. Nature Ecol Evoln 1: 0009

Orchard, M.J., Humphries, S., Schuech, R. & Menden-Deuer, S. (2016) The influence of viscosity on the motility and sensory ability of the dinoflagellate Heterocapsa triquetra. J Plankton Res DOI: 10.1093/plankt/fbw004

Banerji, A., Duncan, A., Griffin, J., Humphries, S., Petchey, O. & Kaltz, O. (2015) Density- and trait-mediated effects of a parasite and a predator in a tri-trophic food web. J Anim Ecol 84(3): 723-733

Ruxton, G.D., Humphries, S., Morrell, L.J. & Wilkinson, D.M. (2014) Why is eusociality an almost exclusively terrestrial phenomenon? J Anim Ecol 83(6): 1248-1255

Hartikainen, H., Humphries, S. & Okamura, B. Form and metabolic scaling in colonial animals. J Exp Biol 217(5): 779-786 (2014)

Simpson, J.L., Humphries, S., Evans, J.P., Simmons, L.W. & Fitzpatrick, J.L. Relationships between sperm length and speed differ among three internally and three externally fertilizing species. Evolution 68(1): 92-104 (2013)

Jeffries, D.L., Chapman, J., Roy, H.E., Humphries, S., Harrington, R., Brown, P.M.J. & Lawson-Handley, L.J. Characteristics and drivers of high-altitude ladybird flight: insights from vertical-looking entomological radar. PLOS ONE 8(12) doi: 10.1371/journal.pone.0082278 (2013)

Humphries, S. A physical explanation of the temperature dependence of physiological processes mediated by cilia and flagella. PNAS 110(36): 14693-14698 (2013)

Wyatt, A.S.J. , Lowe, R.J., Humphries, S. & Waite, A.M. (2013) Particulate nutrient fluxes over a fringing coral reef: Source-sink dynamics inferred from carbon to nitrogen ratios and stable isotopes. Limnol Oceanogr 58(1): 409-427

Wyatt, A.S.J., Waite, A. M. & Humphries, S. Stable isotope analysis reveals community-level variation in fish trophodynamics across a fringing coral reef. Coral Reefs 1–16 (2012).

Anyon, M.J., Orchard, M. J., Buzza, D.M.A., Humphries, S. & Kohonen, M.M. Effect of particulate contamination on adhesive ability and repellence in two species of ant (Hymenoptera; Formicidae). J Exp Biol 215, 605–616 (2012).

Orchard, M.J., Kohonen, M.M. & Humphries, S. The influence of surface energy on the self-cleaning of insect adhesive devices. J Exp Biol 215, 279–286 (2012).

Wyatt, A.S.J., Falter, J.L., Lowe, R.J., Humphries, S. & Waite, A.M. Oceanographic forcing of nutrient uptake and release over a fringing coral reef. Limnol. Oceanogr 57, 401–419 (2012).

Moss, S.P., Joyce, D.A., Humphries, S.., Tindall, K.J. & Lunt, D.H. Comparative analysis of teleost genome sequences reveals an ancient intron size expansion in the zebrafish lineage. Genome Biol Evol doi:10.1093/gbe/evr090 (2011)

Beveridge, O.S., Petchey, O.L. & Humphries, S. Mechanisms of temperature-dependent swimming: the importance of physics, physiology and body size in determining protist swimming speed. J Exp Biol 213, 4223–4231 (2010).

Beveridge, O.S., Petchey, O.L. & Humphries, S. Direct and indirect effects of temperature on the population dynamics and ecosystem functioning of aquatic microbial ecosystems. J Anim Ecol 79, 1324–1331 (2010).

Tomkins, J.L., LeBas, N.R., Witton, M.P., Martill, D.M. & Humphries, S. Positive allometry and the prehistory of sexual selection. Am Nat 176, 141–148 (2010).

Beveridge, O.S., Humphries, S. & Petchey, O.L. The interacting effects of temperature and food chain length on trophic abundance and ecosystem function. J Anim Ecol 79, 693–700 (2010).

Wyatt, A.S.J., Lowe, R.J., Humphries, S. & Waite, A.M. Particulate nutrient fluxes over a fringing coral reef: Relevant scales of phytoplankton Production and mechanisms of supply. Mar Ecol Prog Ser 405, 113–130 (2010).

Wyatt, A.S.J., Waite, A.M. & Humphries, S. Variability in isotope discrimination factors in coral reef fishes: implications for diet and food web reconstruction. PLOS ONE 5, e13682 (2010).

Mossman, J., Slate, J., Humphries, S. & Birkhead, T.R. Sperm morphology and velocity are genetically codetermined in the zebra finch. Evolution 63, 2730–2737 (2009).

Humphries, S. Filter feeders and plankton increase particle encounter rates through flow regime control. PNAS 106, 7882–7887 (2009).

Ruxton, G.D. & Humphries, S. Can ecological and evolutionary arguments solve the riddle of the missing marine insects? Mar Ecol 29, 72–75 (2008).

Humphries, S., Evans, J.P. & Simmons, L.W. Sperm competition: linking form to function. BMC Evol Biol 8, 319 (2008).

Humphries, S., Bonser, R.H.C., Witton, M.P. & Martill, D.M. Did pterosaurs feed by skimming? Physical modelling and anatomical evaluation of an unusual feeding method. PLoS Biol 5, e204 (2007).

Humphries, S. in Body size: the structure and function of aquatic ecosystems (Hildrew, A. G., Raffaelli, D. G. & Edmonds-Brown, R.) 98–117 (Cambridge University Press, 2007).doi:10.1017/CBO9780511611223.003

Brown, S.A., Ruxton, G.D., Pickup, R. & Humphries, S. Seston capture by Hydropsyche siltalai and the accuracy of capture efficiency estimates. Freshwat Biol 50, 113–126 (2005).

Jackson, A.L., Humphries, S. & Ruxton, G.D. Resolving the departures of observed results from the ideal free distribution with simple random movements. J Anim Ecol 73, 612–622 (2004).

Brown, S.A., Ruxton, G.D. & Humphries, S. Physical properties of Hydropsyche siltali (Trichoptera) net silk. J N Am Benthol Soc 23, 771–779 (2004).

Ruxton, G.D. & Humphries, S. Non-IFD movements: reflections on past work and prospects for future developments. Evol Ecol Res (2003).

Humphries, S. & Ruxton, G.D. Estimation of intergenerational drift dispersal distances and mortality risk for aquatic macroinvertebrates. Limnol Oceanogr (2003).

Humphries, S. & Ruxton, G.D. Why did some ichthyosaurs have such large eyes? J Exp Biol 205, 439–441 (2002).

Humphries, S. Dispersal in drift-prone macroinvertebrates: a case for density-independence. Freshwat Biol 47, 921–929 (2002).

Humphries, S. & Ruxton, G.D. Is there really a drift paradox? J Anim Ecol 71, 151–154 (2002).

Ruxton, G.D., Humphries, S. & Farnsworth, K.D. Non‐competitive phenotypic differences can have a strong effect on ideal free distributions. J Anim Ecol 70, 25–32 (2001).

Humphries, S., Ruxton, G.D. & Metcalfe, N.B. Group size and relative competitive ability: geometric progressions as a conceptual tool. Behav Ecol Sociobiol 47, 113–118 (2000).

Humphries, S., Ruxton, G.D. & Metcalfe, N.B. Patch choice and risk: relative competitive ability is context dependent. Anim Behav 58, 1131–1138 (1999).

Ruxton, G.D., Armstrong, J. & Humphries, S. Modelling territorial behaviour of animals in variable environments. Anim Behav 58, 113–120 (1999).

Humphries, S. & Ruxton, G.D. Bower-building: coevolution of display traits in response to the costs of female choice? Ecol Letts 2, 404–413 (1999).

Ruxton, G.D. & Humphries, S. Multiple ideal free distributions of unequal competitors. Evol Ecol Res 1, 635–640 (1999).

Humphries, S., Metcalfe, N.B. & Ruxton, G.D. The effect of group size on relative competitive ability. Oikos (1999).

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